The transformation of agricultural landscapes involves the conversion of forests, scrub, and grasslands into fields, and pastures, as well as the expansion and amalgamation of preexisting fields to enhance farming efficiency (Tscharntke et al., 2005). More recent changes have resulted in more homogeneous agricultural landscapes with greater fragmentation of natural habitats and resulting negative effects on biodiversity. Insects and animals displaced by land-use change must find shelter in remaining fragmented patches of semi-natural land, and those that do not perish (because they do not find alternative areas) must travel further to find appropriate unoccupied habitat and food resources (Marshall and Moonen, 2002). While the trend has been to simplify agricultural production systems, heterogeneous agroecosystems with diverse and connected natural areas are potentially more resilient than simple, homogeneous landscapes (e.g. in adapting to disturbances such as more extreme weather events associated with climate change) (Tscharntke et al., 2005). As a result, the importance of uncropped natural and semi-natural areas in agricultural landscapes is better recognized. For example, the European Union Common Agricultural Policy (CAP) for 2023–2027 requires that at least 4% of eligible land be devoted to biodiversity and landscape protection on all farms of at least 10 ha (European Commission, 2023).