The Neanderthal Mousterian culture of the Ardennes region produced flint tools between 80–35,000 years ago. Omalien tribes of the early Neolithic period (5–4000 BC) developed settled agricultural practices, sophisticated tools and decorated black pottery. The Bronze Age Hilversum culture left evidence of contact overseas in Wessex, southern England. During the pre-Roman period Celtic and Germanic tribes moved across the region. Much of northern Gaul and southern Britain was settled by a Celtic group known as the Belgae, forming numerous tribes including the seafaring Morini and Menapii in what is now Flanders and the bellicose Nervii in Artois. In alliance with Germanic and other Belgic tribes, the Nervii led resistance to the invasion of Julius Caesar in 59 BC, succumbing five years later. Roman control was extended as far north as the Rhine and the area divided into the provinces of Gallia Belgica and Germania Inferior. Several tribes survived as Roman administrative civitates.