The Twa—hunter-gatherer pygmies—were the first people to inhabit Rwanda. They now comprise 1% of the population. The Hutu were the next group to settle in Rwanda. They arrived at some point between AD 500 and 1100. They were small-scale agriculturalists, led by a king who ruled over clan groups. The final group to migrate was the Tutsi around 1400. Their ownership of cattle and their combat skills allowed them to gain control of the country. A feudalistic system developed where the Tutsi lent cows to the Hutu in return for labour and military service. At the apex was the Tutsi king, the mwami (pl., abami), believed to be of divine origin. The abami consolidated their power by centralizing the monarchy and reducing the power of neighbouring chiefs. Mwami Kigeri IV (reigned 1853–95) established the borders of Rwanda in the 19th century.