This study examines the ongoing reconfiguration of heritage engagement through AI, as evidenced by the Digital Dunhuang Project. We conceptualize this shift as an emerging “AI-Enabled Complementary Symbiotic Net-work” comprising five interdependent dimensions—digital immersion, creative co-creation, value co-building, participatory stewardship, and transnational governance—activated through emotion-driven, incentive-based, and technology-mediated mechanisms. Rather than presenting a definitive model, our analysis reveals an evolving architecture where public participation dynamically intersects across these dimensions, with AI serving as both an enabler and mediator. The DDP case demonstrates how such configurations can nurture organic value cycles while accommodating cultural pluralism, suggesting pathways for future development in digital heritage ecosystems.