Morphology of the Shoot System of Scolochloa festucacea (Poaceae)
摘要
The biomorphological features of Scolochloa festucacea (Willd.) Link in the central part of European Russia have been studied; the modular and structural–functional organization of the shoot system has been considered and the life form has been clarified. It is found that S. festucacea is characterized by the innovation of a mixed type (a combination of intra- and extravaginal types) and a combination of silleptic and cataleptic types of branching. The presence of extravaginal renewal is manifested in the potential for the formation of partial bushes, the presence of which contributes to a stronger fixation of the plant in space. The combination of two types of branching leads to heterochrony, the development of buds at different times. The possibility of lodging of shoots, their further rooting in nodes, and the subsequent development of axillary buds into orthotropic shoots has been shown; this contributes to the correction of the spatial structure of the clone, the growth of its total assimilating surface, and a potential increase in reproductive function. A comparative analysis of the modular organization of S. festucacea has shown its similarity in the number and types of elementary modules with those of the long-rhizome sympodial growth of long-shoot helophytes. The analysis of the structural and functional organization of S. festucacea monocarpic shoots has revealed the presence of three variants differing in a set of structural and functional zones.