Grevenius granulifer (Thulin, 1928) revised: a fresh look at one of the most intensively studied water bears (Eutardigrada: Isohypsibioidea)

Redescriptions of species established in the incipient stage of tardigrade taxonomy, in the first half of 20th century, have currently become a routine. Especially type species of all genera should undergo an urgent revision to allow more precise diagnoses and unravel species complexes. Herein, an aquatic species Grevenius granulifer is amended based on the topotypic population from a pond in the Botanical Garden of Lund. Detailed qualitative and quantitative characters alongside a full set of standard DNA barcodes (18S rRNA, 28S rRNA, ITS-2, and COI) are provided. This will open a window for systematic works on other representatives of Grevenius and delivers new data on rarely addressed freshwater tardigrades.