<p>Gestures are an important component of nonverbal communication and are often produced along with speech to aid comprehension, direct attention, and provide emphasis. This study investigates the effects of speaker and addressee visibility on gesture production in computer mediated communication. Based on previous research, we hypothesized that speakers would produce more representational gestures, which convey ideas, actions, and objects, when they could see their addressee’s face and were visible to their addressee, supporting the facial cueing and perspective taking hypotheses. These hypotheses suggest that speakers adjust their gesture use to supplement their speech by considering what their addressee can see (perspective taking) or by using cues from their addressee’s facial expressions (facial cueing). Participants viewed six cartoon clips and retold narrative descriptions of each clip to a confederate via videoconferencing, with visibility conditions varied independently for the speaker and the addressee. Speakers produced significantly more representational gestures when they were visible to the confederate than when they were not. By contrast, speakers’ representational gesture production did not differ when they could see the confederate’s face and when the confederate’s camera was off; however, speakers produced fewer representational gestures when the addressee’s torso was visible than in these conditions. The findings of this study support the perspective taking hypothesis but fail to support the facial cuing hypothesis, providing insight into how gestures are used to communicate in videoconferencing settings.</p>